Author: Purerides.online

  • Cattle 

    Cattle (Bos taurus) are large, domesticatedbovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus Bos. Mature female cattle are called cows and mature male cattle are bulls. Young female cattle are called heifers, young male cattle are oxen or bullocks, and castrated male cattle are known as steers.

    Cattle are commonly raised for meatfor dairy products, and for leather. As draft animals, they pull carts and farm implements. Cattle are considered sacred animals within Hinduism, and it is illegal to kill them in some Indian states. Small breeds such as the miniature Zebu are kept as pets.

    Taurine cattle are widely distributed across Europe and temperate areas of Asia, the Americas, and Australia. Zebus are found mainly in India and tropical areas of Asia, America, and Australia. Sanga cattle are found primarily in sub-Saharan Africa. These types, sometimes classified as separate species or subspecies, are further divided into over 1,000 recognized breeds.

    Around 10,500 years ago, taurine cattle were domesticated from wild aurochs progenitors in central Anatolia, the Levant and Western Iran. A separate domestication event occurred in the Indian subcontinent, which gave rise to zebu. There were over 940 million cattle in the world by 2022. Cattle are responsible for around 7% of global greenhouse gas emissions. They were one of the first domesticated animals to have a fully-mapped genome.

    Etymology

    Further information: List of cattle terminology

    The term cattle was borrowed from Anglo-Norman catel (replacing native Old English terms like kine, now considered archaic, poetic, or dialectal),[1] itself from Medieval Latin capitale ‘principal sum of money, capital’, itself derived in turn from Latin caput ‘head’. Cattle originally meant movable personal property, especially livestock of any kind, as opposed to real property (the land, which also included wild or small free-roaming animals such as chickens—they were sold as part of the land).[2] The word is a variant of chattel (a unit of personal property) and closely related to capital in the economic sense.[3][2] The word cow came via Anglo-Saxon  (plural ), from Common Indo-European gʷōus (genitive gʷowés) ‘a bovine animal’, cf. PersiangâvSanskritgo-.[4] In older English sources such as the King James Version of the Bible, cattle often means livestock, as opposed to deer, which are wild.[2]

    Characteristics

    Description

    Skeleton

    Anatomical model, showing the large 4-chambered stomach

    Cattle are large artiodactylsmammals with cloven hooves, meaning that they walk on two toes, the third and fourth digits. Like all bovid species, they can have horns, which are unbranched and are not shed annually.[5] Coloration varies with breed; common colors are black, white, and red/brown, and some breeds are spotted or have mixed colors.[6] Bulls are larger than cows of the same breed by up to a few hundred kilograms. British Hereford cows, for example, weigh 600–800 kg (1,300–1,800 lb), while the bulls weigh 1,000–1,200 kg (2,200–2,600 lb).[7] Before 1790, beef cattle averaged only 160 kg (350 lb) net. Thereafter, weights climbed steadily.[8][9] Cattle breeds vary widely in size; the tallest and heaviest is the Chianina, where a mature bull may be up to 1.8 m (5 ft 11 in) at the shoulder, and may reach 1,280 kg (2,820 lb) in weight.[10] The natural life of domestic cattle is some 25–30 years. Beef cattle go to slaughter at around 18 months, and dairy cows at about five years.[11]

    Digestive system

    Further information: Digestive system of ruminants

    Bacteria dominate the rumen microbiome; composition can change substantially with diet.[12]

    Cattle are ruminants, meaning their digestive system is highly specialized for processing plant material such as grass rich in cellulose, a tough carbohydrate polymer which many animals cannot digest. They do this in symbiosis with micro-organisms – bacteriafungi, and protozoa – that possess cellulasesenzymes that split cellulose into its constituent sugars. Among the many bacteria that contribute are Fibrobacter succinogenesRuminococcus flavefaciens, and Ruminococcus albus. Cellulolytic fungi include several species of Neocallimastix, while the protozoa include the ciliates Eudiplodinium maggie and Ostracodinium album.[13] If the animal’s feed changes over time, the composition of this microbiome changes in response.[12]

    Cattle have one large stomach with four compartments; the rumenreticulumomasum, and abomasum. The rumen is the largest compartment and it harbours the most important parts of the microbiome.[12] The reticulum, the smallest compartment, is known as the “honeycomb”. The omasum’s main function is to absorb water and nutrients from the digestible feed. The abomasum has a similar function to the human stomach.[14]

    Cattle regurgitate and re-chew their food in the process of chewing the cud, like most ruminants. While feeding, cows swallow their food without chewing; it goes into the rumen for storage. Later, the food is regurgitated to the mouth, a mouthful at a time, where the cud is chewed by the molars, grinding down the coarse vegetation to small particles. The cud is then swallowed again and further digested by the micro-organisms in the cow’s stomach.[14]

    Reproduction

    Nine sequential photos showing the calf being born
    A cow giving birth

    The gestation period for a cow is about nine months long. The ratio of male to female offspring at birth is approximately 52:48.[15] A cow’s udder has two pairs of mammary glands or teats.[16] Farms often use artificial insemination, the artificial deposition of semen in the female’s genital tract; this allows farmers to choose from a wide range of bulls to breed their cattle. Estrus too may be artificially induced to facilitate the process.[17] Copulation lasts several seconds and consists of a single pelvic thrust.[18]

    Cows seek secluded areas for calving.[19] Semi-wild Highland cattle heifers first give birth at 2 or 3 years of age, and the timing of birth is synchronized with increases in natural food quality. Average calving interval is 391 days, and calving mortality within the first year of life is 5%.[20] Beef calves suckle an average of 5 times per day, spending some 46 minutes suckling. There is a diurnal rhythm in suckling, peaking at roughly 6am, 11:30am, and 7pm.[21] Under natural conditions, calves stay with their mother until weaning at 8 to 11 months. Heifer and bull calves are equally attached to their mothers in the first few months of life.[22]

    Cognition

    Individual cattle differ in personality traits such as fearfulness and sociability.[23]

    Cattle have a variety of cognitive abilities. They can memorize the locations of multiple food sources,[24] and can retain memories for at least 48 days.[25] Young cattle learn more quickly than adults,[26] and calves are capable of discrimination learning,[27] distinguishing familiar and unfamiliar animals,[28] and between humans, using faces and other cues.[29] Calves prefer their own mother’s vocalizations to those of an unfamiliar cow.[30] Vocalizations provide information on the age, sex, dominance status and reproductive status of the caller, and may indicate estrus in cows and competitive display in bulls.[31] Cows can categorize images as familiar and unfamiliar individuals.[28] Cloned calves from the same donor form subgroups, suggesting that kin discrimination may be a basis of grouping behaviour.[32] Cattle use visual/brain lateralisation when scanning novel and familiar stimuli.[33] They prefer to view novel stimuli with the left eye (using the right brain hemisphere), but the right eye for familiar stimuli.[34] Individual cattle have also been observed to display different personality traits, such as fearfulness and sociability.[23]

    Senses

    Vision is the dominant sense; cattle obtain almost half of their information visually.[35] Being prey animals, cattle evolved to look out for predators almost all around, with eyes that are on the sides of their head rather than the front. This gives them a field of view of 330°, but limits binocular vision (and therefore stereopsis) to some 30° to 50°, compared to 140° in humans.[28] They are dichromatic, like most mammals.[36] Cattle avoid bitter-tasting foods, selecting sweet foods for energy. Their sensitivity to sour-tasting foods helps them to maintain optimal ruminal pH.[35] They seek out salty foods by taste and smell to maintain their electrolyte balance.[37] Their hearing is better than that of horses,[38] but worse at localising sounds than goats, and much worse than dogs or humans.[39] They can distinguish between live and recorded human speech.[40] Olfaction probably plays a large role in their social life, indicating social and reproductive status.[35][41] Cattle can tell when other animals are stressed by smelling the alarm chemicals in their urine.[42] Cattle can be trained to recognise conspecific individuals using olfaction only.[41]

    Behavior

    Dominance hierarchy

    Spectators in Oman watch a fight between bulls.

    Cattle live in a dominance hierarchy. This is maintained in several ways. Cattle often engage in mock fights where they test each other’s strength in a non-aggressive way. Licking is primarily performed by subordinates and received by dominant animals. Mounting is a playful behavior shown by calves of both sexes and by bulls and sometimes by cows in estrus,[43] however, this is not a dominance related behavior as has been found in other species.[20] Dominance-associated aggressiveness does not correlate with rank position, but is closely related to rank distance between individuals.[20] The horns of cattle are honest signals used in mate selection. Horned cattle attempt to keep greater distances between themselves and have fewer physical interactions than hornless cattle, resulting in more stable social relationships.[44] In calves, agonistic behavior becomes less frequent as space allowance increases, but not as group size changes, whereas in adults, the number of agonistic encounters increases with group size.[45]

    Dominance relationships in semi-wild highland cattle are very firm, with few overt aggressive conflicts: most disputes are settled by agonistic (non-aggressive, competitive) behaviors with no physical contact between opponents, reducing the risk of injury. Dominance status depends on age and sex, with older animals usually dominant to young ones and males dominant to females. Young bulls gain superior dominance status over adult cows when they reach about 2 years of age.[20]

    Grazing behavior

    Charolais bull grazing

    Cattle eat mixed diets, but prefer to eat approximately 70% clover and 30% grass. This preference has a diurnal pattern, with a stronger preference for clover in the morning, and the proportion of grass increasing towards the evening.[46] When grazing, cattle vary several aspects of their bite, i.e. tongue and jaw movements, depending on characteristics of the plant they are eating. Bite area decreases with the density of the plants but increases with their height. Bite area is determined by the sweep of the tongue; in one study observing 750-kilogram (1,650 lb) steers, bite area reached a maximum of approximately 170 cm2 (30 sq in). Bite depth increases with the height of the plants. By adjusting their behavior, cattle obtain heavier bites in swards that are tall and sparse compared with short, dense swards of equal mass/area.[47] Cattle adjust other aspects of their grazing behavior in relation to the available food; foraging velocity decreases and intake rate increases in areas of abundant palatable forage.[48] Cattle avoid grazing areas contaminated by the faeces of other cattle more strongly than they avoid areas contaminated by sheep,[49] but they do not avoid pasture contaminated by rabbits.[50]

    Temperament and emotions

    Ear postures of cows indicate emotional state and overall welfare.[51]

    In cattle, temperament or behavioral disposition can affect productivity, overall health, and reproduction.[52] Five underlying categories of temperament traits have been proposed: shyness–boldness, exploration–avoidance, activity, aggressiveness, and sociability.[53] There are many indicators of emotion in cattle. Holstein–Friesian heifers that had made clear improvements in a learning experiment had higher heart rates, indicating an emotional reaction to their own learning.[54] After separation from their mothers, Holstein calves react, indicating low mood.[55] Similarly, after hot-iron dehorning, calves react to the post-operative pain.[56] The position of the ears has been used as an indicator of emotional state.[28] Cattle can tell when other cattle are stressed by the chemicals in their urine.[42] Cattle are gregarious, and even short-term isolation causes psychological stress. When heifers are isolated, vocalizations, heart rate and plasma cortisol all increase. When visual contact is re-instated, vocalizations rapidly decline; heart rate decreases more rapidly if the returning cattle are familiar to the previously isolated individual.[57] Mirrors have been used to reduce stress in isolated cattle.[58]

    Sleep

    Further information: Sleep in non-human animals

    The average sleep time of a domestic cow is about 4 hours a day.[59] Cattle do have a stay apparatus,[60] but do not sleep standing up;[61] they lie down to sleep deeply.[62]

    Genetics

    Further information: Bovine genome

    Genomic analysis shows there are five main cattle sub-types, here labelled by continent.[63]

    In 2009, the National Institutes of Health and the US Department of Agriculture reported having mapped the bovine genome.[64] Cattle have some 22,000 genes, of which 80% are shared with humans; they have about 1000 genes that they share with dogs and rodents, but not with humans. Using this bovine “HapMap”, researchers can track the differences between breeds that affect meat and milk yields.[65] Early research focused on Hereford genetic sequences; a wider study mapped a further 4.2% of the cattle genome.[63]

    Behavioral traits of cattle can be as heritable as some production traits, and often, the two can be related.[66] The heritability of temperament (response to isolation during handling) has been calculated as 0.36 and 0.46 for habituation to handling.[67] Rangeland assessments show that the heritability of aggressiveness in cattle is around 0.36.[68]

    Quantitative trait loci have been found for a range of production and behavioral characteristics for both dairy and beef cattle.[69]

    Evolution

    Phylogeny

    Cattle have played a key role in human history, having been domesticated since at least the early neolithic age. Archaeozoological and genetic data indicate that cattle were first domesticated from wild aurochs (Bos primigenius) approximately 10,500 years ago. There were two major areas of domestication: one in central Anatolia, the Levant and Western Iran, giving rise to the taurine line, and a second in the area that is now Pakistan, resulting in the indicine line.[70] Modern mitochondrial DNA variation indicates the taurine line may have arisen from as few as 80 aurochs tamed in the upper reaches of Mesopotamia near the villages of Çayönü Tepesi in what is now southeastern Turkey, and Dja’de el-Mughara in what is now northern Syria.[71]

    Although European cattle are largely descended from the taurine lineage, gene flow from African cattle (partially of indicine origin) contributed substantial genomic components to both southern European cattle breeds and their New World descendants.[70] A study on 134 breeds showed that modern taurine cattle originated from Africa, Asia, North and South America, Australia, and Europe.[72] Some researchers have suggested that African taurine cattle are derived from a third independent domestication from the North African aurochs.[70] Whether there have been two or three domestications, European, African, and Asian cattle share much of their genomes both through their species ancestry and through repeated migrations of livestock and genetic material between species, as shown in the diagram.[73]

    Taxonomy

    See also: Bos and Bovinae

    Cattle phylogeny and migrations involve two major species, at least two domestications, and migrations between these.[73]
    Żubroń, a European bison–cattle hybrid

    Cattle were originally identified as three separate species: Bos taurus, the European or “taurine” cattle (including similar types from Africa and Asia); Bos indicus, the Indicine or “zebu”; and the extinct Bos primigenius, the aurochs. The aurochs is ancestral to both zebu and taurine cattle.[74] They were later reclassified as one species, Bos taurus, with the aurochs (B. t. primigenius), zebu (B. t. indicus), and taurine (B. t. taurus) cattle as subspecies.[75] However, this taxonomy is contentious, and authorities such as the American Society of Mammalogists treat these taxa as separate species.[76][77]

    Complicating the matter is the ability of cattle to interbreed with other closely related species. Hybrid individuals and even breeds exist, not only between taurine cattle and zebu (such as the sanga cattle (Bos taurus africanus x Bos indicus), but also between one or both of these and some other members of the genus Bos – yaks (the dzo or yattle[78]), banteng, and gaur. Hybrids such as the beefalo breed can even occur between taurine cattle and either species of bison, leading some authors to consider them part of the genus Bos, as well.[79] The hybrid origin of some types may not be obvious – for example, genetic testing of the Dwarf Lulu breed, the only taurine-type cattle in Nepal, found them to be a mix of taurine cattle, zebu, and yak.[80]

    N’dama cattle in a livestock market in Mali

    The aurochs originally ranged throughout Europe, North Africa, and much of Asia. In historical times, its range became restricted to Europe, and the last known individual died in Mazovia, Poland, around 1627.[81] Breeders have attempted to recreate a similar appearance to the aurochs by crossing traditional types of domesticated cattle, producing the Heck breed.[82]

    A group of taurine-type cattle exist in Africa; they either represent an independent domestication event or were the result of crossing taurines domesticated elsewhere with local aurochs, but they are genetically distinct;[83] some authors name them as a separate subspecies, Bos taurus africanus.[84] The only pure African taurine breeds remaining are the N’DamaKuri and some varieties of the West African Shorthorn.[85]

    Feral cattle are those that have been allowed to go wild.[86] Populations exist in many parts of the world,[87][88] sometimes on small islands.[89] Some, such as Amsterdam Island cattle,[75] Chillingham cattle,[90] and Aleutian wild cattle have become sufficiently distinct to be described as breeds.[91]

    Husbandry

    Practices

    Further information: Animal husbandry

    In concentrated animal feeding operations, the cattle are not allowed to wander and graze, as food is brought to them in a feedlot.[92]

    Cattle are often raised by allowing herds to graze on the grasses of large tracts of rangeland. Raising cattle extensively in this manner allows the use of land that might be unsuitable for growing crops. The most common interactions with cattle involve daily feeding, cleaning and milking. Many routine husbandry practices involve ear taggingdehorning, loading, medical operationsartificial insemination, vaccinations and hoof care, as well as training for agricultural shows and preparations. Around the world, Fulani husbandry rests on behavioural techniques, whereas in Europe, cattle are controlled primarily by physical means, such as fences.[93] Breeders use cattle husbandry to reduce tuberculosis susceptibility by selective breeding and maintaining herd health to avoid concurrent disease.[94]

    In the United States, many cattle are raised intensively, kept in concentrated animal feeding operations, meaning there are at least 700 mature dairy cows or at least 1000 other cattle stabled or confined in a feedlot for “45 days or more in a 12-month period”.[92]

    • A Hereford being inspected for ticks. Cattle are often restrained in cattle crushes when given medical attention.
    • A calf with a nose ring to prevent it from suckling, usually to assist in weaning
    • Cattle feedlot in Colorado, United States

    Population

    Cattle headcounts by country, as of 2021

    Historically, the cattle population of Britain rose from 9.8 million in 1878 to 11.7 million in 1908, but beef consumption rose much faster. Britain became the “stud farm of the world” exporting livestock to countries where there were no indigenous cattle. In 1929 80% of the meat trade of the world was products of what were originally English breeds. There were nearly 70 million cattle in the US by the early 1930s.[95]

    Cattle have the largest biomass of any animal species on Earth, at roughly 400 million tonnes, followed closely by Antarctic krill at 379 million tonnes and humans at 373 million tonnes.[96] In 2023, the countries with the most cattle were India with 307.5 million (32.6% of the total), Brazil with 194.4 million, and China with 101.5 million, out of a total of 942.6 million in the world.[97]

    Economy

    Cattle are kept on farms to produce meat, milk, and leather, and sometimes to pull carts or farm implements.[98]

    Meat

    Further information: Beef cattle and Beef

    The meat of adult cattle is known as beef, and that of calves as veal. Other body parts are used as food products, including blood, liverkidneyheart and oxtail. Approximately 300 million cattle, including dairy animals, are slaughtered each year for food.[99] About a quarter of the world’s meat comes from cattle.[100] World cattle meat production in 2021 was 72.3 million tons.[101]

    • The Hereford is a widespread beef breed, introduced in the 18th century
    • Australian Droughtmaster cattle on an extensive farm in Queensland, Australia
    • Aberdeen Angus, a popular small breed, here in Austria with a traditional cattle bell
    • FAO data for 2021
    • Beef is the third most commonly consumed meat worldwide.
    • Beef (and buffalo meat) production has grown substantially over the recent 60 years.
    • Production of beef worldwide, by country in 2021.

    Dairy

    Main articles: Dairy cattle and Dairy product

    Certain breeds of cattle, such as the Holstein-Friesian, are used to produce milk,[102][103] much of which is processed into dairy products such as buttercheese, and yogurt. Dairy cattle are usually kept on specialized dairy farms designed for milk production. Most cows are milked twice per day, with milk processed at a dairy, which may be onsite at the farm or the milk may be shipped to a dairy plant for eventual sale of a dairy product.[104] Lactation is induced in heifers and spayed cows by a combination of physical and psychological stimulation, by drugs, or by a combination of those methods.[105] For mother cows to continue producing milk, they give birth to one calf per year. If the calf is male, it is generally slaughtered at a young age to produce veal.[106] Cows produce milk until three weeks before birth.[103] Over the last fifty years, dairy farming has become more intensive to increase the yield of milk produced by each cow. The Holstein-Friesian is the breed of dairy cow most common in the UK, Europe and the United States. It has been bred selectively to produce the highest yields of milk of any cow. The average in the UK is around 22 litres per day.[102][103]

    Dairy is a large industry worldwide. In 2023, the 27 European Union countries produced 143 million tons of cow’s milk; the United States 104.1 million tons; and India 99.5 million tons.[107] India further produces 94.4 million tons of buffalo milk,[108] making it (in 2023) the world’s largest milk producer; its dairy industry employs some 80 million people.[109]

    • FAO data for 2021
    • World production of bovine milk (cow + buffalo)

    Draft animals

    Further information: Ox

    Oxen used in traditional ploughing – Karnataka

    Oxen are cattle trained as draft animals. Oxen can pull heavier loads and for a longer period of time than horses.[110] Oxen are used worldwide, especially in developing countries. There are some 11 million draft oxen in sub-Saharan Africa,[111] while in 1998 India had over 65 million oxen.[112] At the start of the 21st century, about half the world’s crop production depended on land preparation by draft animals.[113]

    Hides

    Cattle are not often kept solely for hides, and they are usually a by-product of beef production. Hides are used mainly for leather products such as shoes. In 2012, India was the world’s largest producer of cattle hides.[114] Cattle hides account for around 65% of the world’s leather production.[115][116]

    Health

    Pests and diseases

    Further information: Veterinary medicine

    Drooling due to foot-and-mouth disease

    Infected hoof of the same heifer

    Cattle are subject to pests including arthropod parasites such as ticks (which can in turn transmit diseases caused by bacteria and protozoa),[117] and diseases caused by pathogens including bacteria and viruses. Some viral diseases are spread by insects—i.e. bluetongue disease is spread by midgesPsoroptic mange is a disabling skin condition caused by mitesBovine tuberculosis is caused by a bacterium; it causes disease in humans and in wild animals such as deer and badgers.[118] Foot-and-mouth disease is caused by a virus, affects a range of hoofed livestock and is highly contagious.[119] Bovine spongiform encephalopathy is a neurodegenerative disease spread by a prion, a misfolded brain protein, in contaminated meat.[120] Among the intestinal parasites of cattle are Paramphistomum flukes, affecting the rumen, and hookworms in the small intestine.[121]

    Role of climate change

    Main article: Effects of climate change on livestock

    Most of the top 10 beef-producing countries are likely to see lower production with greater temperatures (left) and heat stress (right).[122]

    Climate change is expected to exacerbate heat stress in cattle, and for longer periods.[123] Heat-stressed cattle may experience accelerated breakdown of adipose tissue by the liver, causing lipidosis.[124] Cattle eat less when heat stressed, resulting in ruminal acidosis, which can lead to laminitis. Cattle can attempt to deal with higher temperatures by panting more often; this rapidly decreases carbon dioxide concentrations at the price of increasing pH, respiratory alkalosis. To deal with this, cattle are forced to shed bicarbonate through urination, at the expense of rumen buffering. These two pathologies can both cause lameness.[124] Another specific risk is mastitis.[124] This worsens as Calliphora blowflies increase in number with continued warming, spreading mastitis-causing bacteria.[125] Ticks too are likely to increase in temperate zones as the climate warms, increasing the risk of tick-borne diseases.[126] Both beef and milk production are likely to experience declines due to climate change.[122][127]

    Impact of cattle husbandry

    On public health

    Cattle health is at once a veterinary issue (for animal welfare and productivity), a public health issue (to limit the spread of disease), and a food safety issue (to ensure meat and dairy products are safe to eat). These concerns are reflected in farming regulations.[128] These rules can become political matters, as when it was proposed in the UK in 2011 that milk from tuberculosis-infected cattle should be allowed to enter the food chain.[129] Cattle disease attracted attention in the 1980s and 1990s when bovine spongiform encephalopathy (mad cow disease) broke out in the United Kingdom. BSE can cross into humans as the deadly variant Creutzfeldt–Jakob disease; 178 people in the UK had died from it by 2010.[130]

    On the environment

    Main article: Environmental impact of cattle production

    Beef has the highest greenhouse gas impact of any agricultural commodity, followed by mutton.

    The gut flora of cattle produce methane, a powerful[131] greenhouse gas, as a byproduct of enteric fermentation, with each cow belching out 100kg a year.[132] Additional methane is produced by anaerobic fermentation of stored manure.[133] The FAO estimates that in 2015 around 7% of global greenhouse gas emissions were due to cattle, but this is uncertain.[134] Reducing methane emissions quickly helps limit climate change.[134]

    Concentrated animal feeding operations in particular produce substantial amounts of wastewater and manure,[135][136] which can cause environmental harms such as soil erosion, human and animal exposure to toxic chemicals, development of antibiotic resistant bacteria and an increase in E. coli contamination.[137][138]

    In many world regions, overgrazing by cattle has reduced biodiversity of the grazed plants and of animals at different trophic levels in the ecosystem.[139] A well documented consequence of overgrazing is woody plant encroachment in rangelands, which significantly reduces the carrying capacity of the land over time.[140]

    On animal welfare

    Further information: Cruelty to animals § Welfare concerns of farm animals

    Confining calves for veal production in individual crates has attracted welfare concerns.

    Cattle husbandry practices including branding,[141] castration,[142] dehorning,[143] ear tagging,[144] nose ringing,[145] restraint,[146] tail docking,[147] the use of veal crates,[148] and cattle prods[149] have raised welfare concerns.[150]

    Stocking density is the number of animals within a specified area. High stocking density can affect cattle health, welfare, productivity,[151] and feeding behaviour.[152] Densely-stocked cattle feed more rapidly and lie down sooner, increasing the risk of teat infection, mastitis, and embryo loss.[153][154] The stress and negative health impacts induced by high stocking density such as in concentrated animal feeding operations or feedlots, auctions, and transport may be detrimental to cattle welfare.[155]

    To produce milk, most calves are separated from their mothers soon after birth and fed milk replacement in order to retain the cows’ milk for human consumption.[156]Dairy cattle are frequently artificially inseminated.[157] Animal welfare advocates are critical of this practice, stating that this breaks the natural bond between the mother and her calf.[156] The welfare of veal calves is also a concern.[158]

    Two sports involving cattle are thought to be cruel by animal welfare groups: rodeos and bullfighting. Such groups oppose rodeo activities including bull ridingcalf roping and steer roping, stating that rodeos are unnecessary and cause stress, injury, and death to the animals.[159] In Spain, the Running of the bulls faces opposition due to the stress and injuries incurred by the bulls during the event.[160]

    In culture

    From early in civilisation, cattle have been used in barter.[161][162] Cattle play a part in several religions. Veneration of the cow is a symbol of Hindu community identity.[163] Slaughter of cows is forbidden by law in several states of the Indian Union.[164]

    The ox is one of the 12-year cycle of animals which appear in the Chinese zodiac. The astrological sign Taurus is represented as a bull in the Western zodiac.[165]

  • Goat 

    The goat or domestic goat (Capra hircus) is a species of goat-antelope that is mostly kept as livestock. It was domesticated from the wild goat (C. aegagrus) of Southwest Asia and Eastern Europe. The goat is a member of the family Bovidae, meaning it is closely related to the sheep. It was one of the first animals to be domesticated, in Iran around 10,000 years ago.

    Goats have been used for milkmeatwool, and skins across much of the world. Milk from goats is often turned into cheese. In 2022, there were more than 1.1 billion goats living in the world, of which 150 million were in India.

    Goats feature in mythology, folklore, and religion in many parts of the world, including in the classical myth of Amalthea, in the goats that pulled the chariot of the Norse god Thor, in the Scandinavian Yule goat, and in Hinduism’s goat-headed Daksha. In Christianity and Satanism, the devil is sometimes depicted as a goat.

    Etymology

    The Modern English word goat comes from Old English gāt “goat, she-goat”, which in turn derives from Proto-Germanic *gaitaz (cf. Dutch/Frisian/Icelandic/Norwegian geitGerman Geiß, and Gothic gaits), ultimately from Proto-Indo-European *ǵʰaidos meaning “young goat” (cf. Latin haedus “kid”).[1] To refer to the male goat, Old English used bucca (cf. Dutch/Frisian bok, modern English buck) until ousted by hegotehegoote (‘he-goat’) in the late 12th century.[2] Nanny goat (adult female) originated in the 18th century, and billy goat (adult male) in the 19th century.[3][4]

    Castrated males are called wethers. While the words hircine and caprine both refer to anything having a goat-like quality, hircine is used most often to emphasize the distinct smell of domestic goats.[5][6]

    History

    Further information: Domestication of the goat

    Horn cores from the Neolithic village of Atlit Yam, between 6900 and 6300 BC

    Goats are among the earliest animals to have been domesticated by humans.[7] A genetic analysis[8] confirms the archaeological evidence that the wild bezoar ibex, found today in the Zagros Mountains, but formerly widespread in Anatolia, is the likely original ancestor of all or most domestic goats today.[7]

    Neolithic farmers began to herd wild goats primarily for easy access to milk and meat, as well as to their dung, which was used as fuel; and their bones, hair, and sinew were used for clothing, building, and tools.[9] The earliest remnants of domesticated goats dating 10,000 years Before Present are found in Ganj Dareh in Iran.[10][11] Goat remains have been found at archaeological sites in JerichoChoga Mami,[12] Djeitun, and Çayönü, dating the domestication of goats in Western Asia at between 8,000 and 9,000 years ago.[7] DNA evidence suggests that goats were domesticated around 10,000 years ago.[8] Historically, goat hide has been used for water and wine bottles in both traveling and transporting wine for sale, and to produce parchment.[13]

    Biology

    Description

    Each breed of goat has specific weight ranges, which vary from more than 140 kg (310 lb) for bucks of larger breeds such as the Boer, to 20 to 27 kg (44 to 60 lb) for smaller does.[14] Within each breed, different strains or bloodlines may have different recognized sizes. At the bottom of the size range are miniature breeds such as the African Pygmy, which stand 41 to 58 cm (16 to 23 in) at the shoulder as adults.[15]

    Most goats naturally have two horns, their shape and size depending on the breed.[16] There have been incidents of polycerate goats (having as many as eight horns), although this is a genetic rarity. Unlike cattle, goats have not been successfully bred to be reliably polled, as the genes determining sex and those determining horns are closely linked. Breeding together two genetically polled goats results in a high number of intersex individuals among the offspring, which are typically sterile.[16] Their horns are made of living bone surrounded by keratin and other proteins, and are used for defense, dominance, territoriality,[17] and thermoregulation.[18] Both male and female goats may have beards, and many types of goat (most commonly dairy goats, dairy-cross Boers, and pygmy goats) may have wattles, one dangling from each side of the neck.[19] Goats have horizontal, slit-shaped pupils, allowing them to see well by both night and day, and giving them a wide field of vision on either side to detect predators, while avoiding being dazzled by sunlight from above.[20] Goats have no tear ducts.[21]

    Goats are ruminants. They have a four-chambered stomach consisting of the rumen, the reticulum, the omasum, and the abomasum. As with other mammal ruminants, they are even-toed ungulates. The females have an udder consisting of two teats, in contrast to cattle, which have four teats.[22] An exception to this is the Boer goat, which sometimes may have up to eight teats.[23][24] Goats are diploid with two sets of 30 chromosomes.[25]

    • Skeleton
    • Eye with horizontal pupil

    Comparison with sheep

    Sheep and goats are closely related: both are in the subfamily Caprinae. However, they are separate species, so hybrids rarely occur and are always infertile. A hybrid of a ewe and a buck is called a sheep-goat hybrid. Visual differences between sheep and goats include the beard of goats and the divided upper lip of sheep. Sheep tails hang down, even when short or docked, while the short tails of goats are held upwards. Sheep breeds are often naturally polled (either in both sexes or just in the female), while naturally polled goats are rare (though many are polled artificially). Males of the two species differ in that buck goats acquire a unique and strong odor during the rut, whereas rams do not.[26]

    Behavior and ecology

    Goats are naturally curious. They are agile and able to climb and balance in precarious places. This makes them the only ruminant to regularly climb trees. These behaviours have made them notorious for escaping their pens by testing fences and enclosures. If any of the fencing can be overcome, goats almost inevitably escape. Goats are as intelligent as dogs by some studies.[27] When handled as a group, goats display less herding behavior than sheep. When grazing undisturbed, they spread across the field or range, rather than feed side by side as do sheep. When nursing young, goats leave their kids separated (“lying out”) rather than clumped, as do sheep. They generally turn and face an intruder, and bucks are more likely to charge or butt at humans than are rams.[28] A 2016 study reports that goats try to communicate with people like domesticated animals such as dogs and horses. They look to a human for assistance when faced with a newly-modified challenge.[29][30]

    • Goats grazing in an argan tree, Morocco
    • Goats establishing a dominance hierarchy through head butting
    • Herd browsing together in Japan
    • Moving a herd on a road in Ladakh

    Reproduction

    Goats reach puberty between three and 15 months of age, depending on breed and nutritional status. Many breeders prefer to postpone breeding until the doe has reached 70% of the adult weight, but this separation is rarely possible in extensively managed, open-range herds.[31]

    Bucks (uncastrated males) of Swiss and northern breeds come into rut in the fall as with the does’ heat cycles. Bucks of equatorial breeds may show seasonal reduced fertility, but as with the does, are capable of breeding at all times. Rut is characterized by a decrease in appetite and obsessive interest in the does.[17] A buck in rut displays flehmen lip curling and urinates on his forelegs and face.[32] Sebaceous scent glands at the base of the horns add to the male goat’s odor, which is important to make him attractive to the female. Some does will not mate with a buck which has had its scent glands removed.[17]

    Gestation length is approximately 150 days. Twins are the usual result, with single and triplet births also common. Less frequent are litters of quadrupletquintuplet, and even sextuplet kids. Birthing, known as kidding, generally occurs uneventfully. Just before kidding, the doe will have a sunken area around the tail and hip, as well as heavy breathing. She may have a worried look, become restless and display great affection for her keeper. The mother often eats the placenta, which gives her much-needed nutrients, helps stanch her bleeding, and parallels the behavior of wild herbivores, such as deer, to reduce the lure of the birth scent for predators.[33][34]

    Freshening (coming into milk production) usually occurs at kidding, although milk production is also relatively common in unbred doelings of dairy breeds.[35] Milk production varies with the breed, age, quality, and diet of the doe; dairy goats generally produce between 680 and 1,810 kg (1,500 and 4,000 lb) of milk per 305-day lactation. On average, a good quality dairy doe will give at least 3 kg (6 lb) of milk per day while she is in milk. A first-time milker may produce less, or as much as 7 kg (16 lb), or more of milk in exceptional cases. After the lactation, the doe will “dry off”, typically after she has been bred. Occasionally, goats that have not been bred and are continuously milked will continue lactation beyond the typical 305 days.[36] Male lactation sometimes occurs in goats.[37]

    • Female suckling two kids
    • A two-month-old kid in a field of capeweed
    • A female and two kids

    Diet

    Goats are reputed to be willing to eat almost anything. They are browsing animals, not grazers like cattle and sheep, and (coupled with their highly curious nature) will chew on and taste anything resembling plant matter to decide whether it is good to eat, including cardboard, clothing and paper.[38]

    The digestive physiology of a very young kid (like the young of other ruminants) is essentially the same as that of a monogastric animal. Milk digestion begins in the abomasum, the milk having bypassed the rumen via closure of the reticuloesophageal groove during suckling. At birth, the rumen is undeveloped, but as the kid begins to consume solid feed, the rumen soon increases in size and in its capacity to absorb nutrients.[39]

    The adult size of a particular goat is a product of its breed (genetic potential) and its diet while growing (nutritional potential). As with all livestock, increased protein diets (10 to 14%) and sufficient calories during the prepuberty period yield higher growth rates and larger eventual size than lower protein rates and limited calories.[40] Large-framed goats, with a greater skeletal size, reach mature weight at a later age (36 to 42 months) than small-framed goats (18 to 24 months) if both are fed to their full potential. Large-framed goats need more calories than small-framed goats for maintenance of daily functions.[41]

    • A goat tied to restrict its grazing area
    • A goat feeding in a field of capeweed, toxic to most stock animals

    Diseases and life expectancy

    Main article: List of infectious sheep and goat diseases

    While goats are hardy animals and often need little medical care, they are subject to a number of diseases. Among the conditions affecting goats are respiratory diseases including pneumonia, foot rot, internal parasites, pregnancy toxicosis, and feed toxicity. Goats can become infected with various viral and bacterial diseases, such as foot-and-mouth diseasecaprine arthritis encephalitiscaseous lymphadenitis, pinkeye, mastitis, and pseudorabies. They can transmit a number of zoonotic diseases to people, such as tuberculosisbrucellosisQ fever, and rabies.[42]

    Life expectancy for goats is between 15 and 18 years.[43] An instance of a goat reaching the age of 24 has been reported.[44] Several factors can reduce this average expectancy; problems during kidding can lower a doe’s expected life span to 10 or 11, and stresses of going into rut can lower a buck’s expected life span to eight to 10 years.[44]

    Agriculture

    Husbandry

    Main article: Goat farming

    Further information: List of goat breeds

    Husbandry, or animal care and use, varies by region and culture. The minimal requirements for goats include a grazing area or the bringing of fodder to penned animals, with enough hayracks for all of them to feed simultaneously; fresh water; salt licks; space for the animals to exercise; and disposal of soiled bedding.[45]

    In Africa and the Middle East, goats are typically run in flocks with sheep. This maximizes the production per acre, as goats and sheep prefer different food plants. Multiple types of goat-raising are found in Ethiopia, where four main types have been identified: pastured in annual crop systems, in perennial crop systems, with cattle, and in arid areas, under pastoral (nomadic) herding systems. In all four systems, however, goats were typically kept in extensive systems, with few purchased inputs.[46]

    In Nigeria and in parts of Latin America, some goats are allowed to wander the homestead or village, while others are kept penned and fed in a ‘cut-and-carry’ system. This involves cutting grasses, maize or cane for feed rather than allowing the animal access to the field. The system is well suited for crops like maize that are sensitive to trampling.[47]

    • Goat husbandry in Chile
    • Small-scale goat husbandry in Germany
    • goatherd leading his goats on a rough hillside in Spain
    • smallholder with goats in Burkina Faso
    • Mixed herd of goats and sheep for efficient grazing, Syria[46]

    Worldwide population

    Worldwide distribution of goats in 2015

    The top producers of goat milk in 2022 were India (6.25 million metric tons), Bangladesh (0.91 million metric tons), and South Sudan (0.52 million metric tons).[48] As of 2015, India slaughters 41% of 124.4 million goats each year. The 0.6 million metric tonnes of goat meat make up 8% of India’s annual meat production.[49] Approximately 440 million goats are slaughtered each year for meat worldwide, yielding 6.37 million metric tons of meat.[50]

    Feral goats

    Main article: Feral goat

    Feral goats in a churchyard near Llandudno, Wales

    Goats readily revert to the wild (become feral) if given the opportunity.[7] Feral goats have established themselves in many areas: they occur in Australia, New Zealand, Great Britain, the Galapagos and many other places. When feral goats reach large populations in habitats that provide unlimited water supply and do not contain sufficient large predators or are otherwise vulnerable to goats’ aggressive grazing habits, they may have serious effects, such as removing native scrub and trees. Feral goats are extremely common in Australia, with an estimated 2.6 million in the mid-1990s.[51]

    Uses

    Goats are used to provide milk and specialty wools, and as meat and goatskin.[52][53] Some charities provide goats to impoverished people in poor countries, in the belief that having useful things alleviates poverty better than cash. The cost of obtaining goats and then distributing them can however be high.[54]

    Meat

    Main article: Goat meat

    See also: List of goat dishes

    The taste of goat kid meat is similar to that of spring lamb meat;[55] in fact, in the English-speaking islands of the Caribbean, and in South Asia, the word ‘mutton‘ denotes both goat and sheep meat.[56][57] However, some compare the taste of goat meat to veal or venison, depending on the age and condition of the goat. Its flavor is said to be primarily linked to the presence of 4-methyloctanoic and 4-methylnonanoic acid.[58] The meat is made into dishes such as goat curry,[59] mutton satay,[60] and capra e fagioli.[61]

    Milk, butter, and cheese

    Main article: Goat milk

    See also: List of goat milk cheeses

    Goats produce about 2% of the world’s total annual milk supply.[62] Dairy goats produce an average of 540 to 1,180 kg (1,200 to 2,600 lb) of milk during an average 284-day lactation.[63] The milk can contain between around 3.5% and 5% butterfat according to breed.[64] Goat milk is processed into products including cheese[65] and Dulce de leche.[66]

    Mohair and cashmere wool

    Further information: Mohair and Cashmere wool

    Most goats have soft insulating hairs nearer the skin, and long guard hairs on the surface. The soft hairs are the ones valued by the textile industry; the material goes by names such as down, cashmere and pashmina. The coarse guard hairs are of little value as they are too coarse, difficult to spin and to dye. The cashmere goat produces a commercial quantity of fine and soft cashmere wool, one of the most expensive natural fibers commercially produced. It is harvested once a year.[67] The Angora breed of goats produces long, curling, lustrous locks of mohair. The entire body of the goat is covered with mohair and there are no guard hairs. The locks constantly grow to four inches or more in length. Angora crossbreeds, such as the pygora and the nigora, have been created to produce mohair and/or cashgora on a smaller, easier-to-manage animal. The wool is shorn twice a year, with an average yield of about 4.5 kg (10 lb).[68]

    Land clearing

    Further information: Conservation grazing

    Goats have been used by humans to clear unwanted vegetation for centuries. They have been described as “eating machines” and “biological control agents”.[69][70] There has been a resurgence of this in North America since 1990, when herds were used to clear dry brush from California hillsides thought to be endangered by potential wildfires. This form of using goats to clear land is sometimes known as conservation grazing. Since then, numerous public and private agencies have hired private herds from companies such as Rent A Goat to perform similar tasks.[69][71] This may be expensive and their smell may be a nuisance.[72] This practice has become popular in the Pacific Northwest, where they are used to remove invasive species not easily removed by humans, including (thorned) blackberry vines and poison oak.[69][73][74] Chattanooga, TN and Spartanburg, SC have used goats to control kudzu, an invasive plant species prevalent in the southeastern United States.[75]

    Medical training

    Some countries’ militaries use goats to train combat medics. In the United States, goats have become the main animal species used for this purpose after the Pentagon phased out using dogs for medical training in the 1980s.[76] While modern mannequins used in medical training are quite efficient in simulating the behavior of a human body, trainees feel that “the goat exercise provide[s] a sense of urgency that only real life trauma can provide”. The practice has elicited outcry from animal-rights groups.[77]

    Pets

    Some people choose goats as a pet because of their ability to form close bonds with their human guardians.[78][79] Goats are social animals and usually prefer the company of other goats, but because of their herd mentality, they will follow their owner and form close bonds with them, hence their continuing popularity.[29]

    Goats are similar to deer with regard to nutrition and need a wide range of food, including things like haygrain feed or pelleted grain mix, and loose minerals.[80] Goats generally either inherit certain feeding preferences or learn them after birth.[81]

    In culture

    Mythology, folklore and astrology

    In classical mythAmalthea is either a nymph who fed the infant god Jupiter with goat’s milk, or the goat who suckled the infant. In another legend, the god broke one of the goat’s horns, endowing it with the power to fill itself with whatever its owner wanted, making it the cornucopia or horn of plenty.[82] The ancient city of Ebla in Syria contains a tomb with a throne decorated with bronze goat heads, now called “The Tomb of the Lord of the Goats”.[83][84]

    According to Norse mythology, the god of thunder, Thor, has a chariot that is pulled by the goats Tanngrisnir and Tanngnjóstr At night when he sets up camp, Thor eats the meat of the goats, but takes care that all bones remain whole. Then he wraps the remains up, and in the morning, the goats always come back to life to pull the chariot. When a farmer’s son who is invited to share the meal breaks one of the goats’ leg bones to suck the marrow, the animal’s leg remains broken in the morning, and the boy is forced to serve Thor as a servant to compensate for the damage.[85] Possibly related, the Yule goat (Julbocken) is a Scandinavian Christmas tradition. It originally denoted the goat that was slaughtered around Yule, now more often a goat figure made out of straw. It is used for the custom of going door-to-door singing carols and getting food and drinks in return, often fruit, cakes and sweets. The Gävle goat is a giant version of the yule goat, erected every year in the Swedish city of Gävle.[86][87] In Finland the tradition of NuutinpäiväSt. Knut’s Day, January 13—involves young men dressed as goats (Finnish: Nuuttipukki) who visit houses. Usually the dress was an inverted fur jacket, a leather or birch bark mask, and horns. Unlike the analogous Santa Claus, Nuuttipukki was a scary character (cf. Krampus). The men dressed as Nuuttipukki wandered from house to house, came in, and typically demanded food from the household and especially leftover alcohol. In Finland the Nuuttipukki tradition is kept alive in areas of SatakuntaSouthwest Finland and Ostrobothnia. Nowadays the character is usually played by children and involves a happy encounter.[88]

    The goat is one of the 12-year cycle of animals which appear in the Chinese zodiac.[89] Several mythological hybrid creatures contain goat parts, including the Chimera.[90] The Capricorn constellation sign in the Western zodiac is usually depicted as a goat with a fish’s tail.[91] Fauns and satyrs are mythological creatures with human bodies and goats’ legs.[92] The lustful Greek god Pan similarly has the upper body of a man and the horns and lower body of a goat.[85] A goatee is a tuft of facial hair on a man’s chin, named for its resemblance to a goat’s beard.[93]

    Religion

    In Hinduism, Daksha, one of the prajapati, is sometimes depicted with the head of a male goat. A legend states that Daksha failed to invite Shiva to a sacrifice; Shiva beheaded Daksha, but when asked by Vishnu, restored Daksha to life with the head of a goat.[94] Goats are mentioned many times in the Bible. Their importance in ancient Israel is indicated by the seven different Hebrew and three Greek terms used in the Bible.[95] A goat is considered a “clean” animal by Jewish dietary laws and a kid was slaughtered for an honored guest. It was also acceptable for some kinds of sacrifices. Goat-hair curtains were used in the tent that contained the tabernacle (Exodus 25:4). Its horns can be used instead of sheep’s horn to make a shofar.[96] On Yom Kippur, the festival of the Day of Atonement, two goats were chosen and lots were drawn for them. One was sacrificed and the other allowed to escape into the wilderness, symbolically carrying with it the sins of the community. From this comes the word “scapegoat“.[97] In Matthew 25:31–46, Jesus said that like a shepherd he will separate the nations, rewarding the sheep, those who have shown kindness, but punishing the goats.[95] The devil is sometimes depicted, like Baphomet, as a goat, making the animal a significant symbol throughout Satanism. The inverted pentagram of Satanism is sometimes depicted with a goat’s head of Baphomet, which originated from the Church of Satan.[98]